Using an ensemble of wintertime hindcasts with a high-resolution (T106L60) Atmospheric General Circulation Model (AGCM) forced by observed sea surface temperatures (SSTs) and extending into the stratosphere, we investigate the formation and lifecycle of the Aleutian-Icelandic low Seesaw (AIS) during the 1978 to 1993 period. The AIS has been newly proposed to be an important mode of variability, linking the major wintertime surface lows, the Icelandic Low and the Aleutian Low, in late winter, and thereby linking climate variability over the North Pacific and the North Atlantic. We demonstrate for the first time with a stratosphere-troposphere model, that a coherent, ensemble-mean AIS extension into the stratosphere exists, where its presence modulates ultra-long planetary wave propagation and the polar night jet intensity. The model AIS peaks in February, when the Aleutian and Icelandic Low anti-correlation maximizes at -0.59. The AIS provides a new way to describe the El Niño-Southern Oscillation (ENSO) phenomenon influence into the stratosphere. For example, El-Nino conditions correspond to a deeper than normal Aleutian Low, extending its influence into the Icelandic sector as an AIS negative phase (weakened Icelandic Low), hence enhanced planetary wave vertical propagation and a weakened stratospheric polar vortex. This maturation of the AIS in late winter explains the intra-seasonal variability of the stratospheric response to ENSO, which peaks in late winter. Internal model variability is large and enhanced potential predictability is found primarily in the western hemisphere, with a western Atlantic maxima being more pronounced in the stratosphere than in the upper troposphere.